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What is a costard?

What is a costard? Here are some definitions.

Noun
  1. (Britain) a large cooking apple
  2. the tree of this fruit
  3. (humorous) head
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Examples
The cull thought to have loped by breaking through the crackmans, but we fetched him back by a nope on the costard, which stopped his jaw.
The cooking equivalent was the Costard, which gave rise to the term costermonger, first applied to apple salesmen.

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