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What are holdfasts?

What are holdfasts? Here are some definitions.

Noun
  1. plural of holdfast
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Crowns commonly have some length of column still attached, and holdfasts, less commonly recognized than crowns, also have some length of column attached.
Both of the above forms were found on the washed-up holdfasts of Macrocystis.
Heavy grazing may damage and weaken the holdfasts which anchor the algae.
Sphenothallus may have been equally happy in normal marine conditions, where traces of holdfasts have also been observed.
Found attached to stones, etc., about the holdfasts of fucus.
From the holdfasts rise stalks, and from them grow fronds buoyant with carbon dioxide-filled bubbles called pneumatocysts.

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